ROLE OF SIZE IN MATE FORMATION OF
AMPHIPRION
OCELLARIS UNDER CULTURE
By
Danny C. Chen
A paper submitted in partial
fulfillment of the requirements of the Honors Program Department of Biology and
Marine Biology.
Approved By:
__________________________
Ileana Clavijo, PhD
Faculty
Supervisor
Examining Committee:
__________________________
__________________________
__________________________
__________________________
Department
Chair
_________________________________
Honors Council
Representative
_________________________________
Director of the Honors
Scholars Program
December 2006
Abstract
Individuals of the false clown anemonefish (Amphiprion ocellaris) were placed into groups of different
size. The objectives of this experiment
were to examine the effects of size on mate formation in social groups held in aquaria
and to examine how growth rates were regulated to maintain the variation in
size. Sex is determined by size and
dominance in Amphiprion ocellaris. The largest individual of a group becomes the
rank 1 female. The second largest is a
functional male in rank 2. All others
are subordinate males in competition for the functional male position. Five tanks were set up each containing five
individuals of different sizes. It was
expected that once the top two fish in each tank established a bond, subordinates
would compete for the male position if the top male was removed. However, low density and group behavioral
instability drove the dominant pairs in three of the five tanks to act
aggressively toward the subordinates.
Over time, most subordinates were eliminated. Fish left in each tank were measured and
growth rates were calculated. The most
dominant fish (females) in social groups persisted and had the greatest mean
growth rates of 0.898, 1.091, 0.941, 0.941, and 1.011mm/10-d respectively. The original males that bonded with the
females also persisted and generally had the second greatest mean growth rates
of 0.500, 0.984, and 0.672mm/10-d for three social groups. In conclusion, size is important in mate formation. Dominant individuals became dominant by
having a large size advantage; to remain as dominant fish, growth rates of
subordinates were regulated indirectly by dominant fish so that they were
unable to acquire higher ranks.